IGCP 575: Pennsylvanian terrestrial habitats and biotas of Southeastern Euramerica


Dr. Chris J. Cleal
Curriculum Vitae


I started work on late Carboniferous palaeobotany in 1972, studying for a PhD with Bob Wagner at the University of Sheffield. My initial interests were a combination of biostratigraphy and the taxonomy of the Medullosales, based mainly around the Saar-Lorraine floras, and these have stayed with me most of my working life. After leaving Sheffield in 1975, I spent two years in the Geology Department of the National Museum of Wales in Cardiff, where I helped curate the Emily Dix and other collections of Carboniferous plant fossils held there. I returned to Sheffield for a year to do more work with Bob Wagner, and then in 1978 joined the Nature Conservancy Council (NCC), whose geological offices were based in Newbury, Berkshire (where I still live).

The NCC was then undertaking a complete review of the protected geological sites in Britain, and I was responsible for the upper Carboniferous stratigraphical and all palaeobotanical sites. I was required to visit all sites in Britain showing these interests, to assess their scientific importance in a national and international context, ultimately selecting the most important as proposed Sites of Special Scientific Interest. I also was responsible for organising excavations and site cleaning work, which included digs on dinosaur sites on the Isle of Wight, Devonian fish sites in Shropshire, and Carboniferous palaeobotany sites in the Forest of Dean, north Wales, Lancashire and Berwickshire.

The NCC work finished in 1988 and I spent two years as a ‘house husband’ caring for our eldest daughter, Helen. However, in 1990 I was invited by Barry Thomas to join the Botany Department of the National Museum of Wales, where I have been ever since. Initially I acted as a research assistant to Barry but, when the department was later merged with Zoology (to for the Department of Biodiversity & Systematic Biology) I was appointed Assistant Keeper in charge of Vegetation History. During this time I have continued to develop my interest in Carboniferous palaeobotany and terrestrial environments, including the development of a number of international collaborative projects, of which IGC 575 is the most recent.

Current research interests
Taxonomy of Palaeozoic Medullosales, Marattiales and Sphenophyta
Biostratigraphy, biogeography and biodiversity of Palaeozoic floras
The interaction of climate and vegetation in Palaeozoic times
Palaeobotanical taxonomic nomenclature
The history of palaeobotany in Britain


Projects and organisations

Late Variscan terrestrial biotas and palaeoenvironments (IGCP 469): Senior Co-ordinator
Climatic and vegetational changes in the Late Carboniferous tropical belt (Supported by NATO Science Programme): Project Co-ordinator
Bibliography of European Palaeobotany and Palynology: co-editor
International Association for Plant Taxonomy (IAPT), Fossil Plant Committee
IUGS Subcommission on Carboniferous Stratigraphy: Corresponding Member
European Association for the Conservation of Earth Heritage (ProGeo): British Delegate
British Institute for Geological Conservation: Secretary
Fellow of the Linnean Society and the Geological Society, London
Former editor and Council Member of the Palaeontological Association

Publications (Books)
Cleal, C. J (ed.) 1991. Plant fossils in geological investigation: the Palaeozoic. Ellis Horwood, Chichester, 233 pp.

Cleal, C. J. & Thomas, B. A. 1995. Palaeozoic palaeobotany of Great Britain. Chapman & Hall, London, Geological Conservation Review Series, No. 9, xii + 295 pp.

Cleal, C. J. & Thomas, B. A. 1995. British Upper Carboniferous stratigraphy. Chapman & Hall, London, xii + 295 pp. (Geological Conservation Review Series, No. 11)

Cleal, C. J. & Thomas, B. A. 1994. Plant fossils of the British Coal Measures. Palaeontological Association, London, 222 pp. (Field Guide to Fossils).

Cleal, C. J. & Thomas, B. A. 1999. Plant fossils. Boydell Press, Woodbridge, ix + 188 pp., 128 pls. (Fossils Illustrated, Volume 2).

Cleal, C. J., Thomas, B. A., Batten, D. J. & Collinson, M. E. 2001. Mesozoic and Tertiary palaeobotany of Great Britain. Joint Nature Conservation Committee, Peterborough, xviii + 335 pp. (Geological Conservation Review Series, No. 22)

Anderson, J. M., Anderson, H. M. & Cleal, C. J. 2007. Brief history of the gymnosperms: classification, biodiversity, phytogeography and ecology. South Africa National Biodiversity Institute, Pretoria, x + 279 pp (Strelitzia, 20).

Cleal, C. J. & Thomas, B. A. 2009. Introduction to plant fossils. Cambridge University Press, Cambridge, viii +237 pp.

Selected recent papers
Thomas, B. A. & Cleal, C. J. 2001. A new early Westphalian D flora from Aberdulais Falls, South Wales. Proceedings of the Geologists' Association, 112, 373-377.

Šimunek, Z. & Cleal, C. J. 2002. The taxonomy of middle Westphalian medullosalean foliage known as 'Alethopteris grandinioides early form' from the Central and Western Bohemian Basin. Bulletin of the Czech Geological Survey, 77, 61-64.

Shute, C. H. & Cleal, C. J. 2002. Ecology and growth habit of Laveineopteris: a gymnosperm from the Late Carboniferous tropical rain forests. Palaeontology, 45, 943-972.

Cleal, C. J. & Wang Ziqiang. 2002. A new and diverse plant fossil assemblage from the upper Westphalian Benxi Formation, Shanxi, China, and its palaeofloristic significance. Geological Magazine, 139, 107-130.

Cleal, C. J. 2002. Two new late Carboniferous Neuropteris species (Medullosales) from Saarland, Germany and their palaeogeographical significance. Botanical Journal of the Linnean Society, 139, 193-205.

Evans, B. G., Cleal, C. J., Thomas, B. A. & Wimbledon, W. A. P. 2003. The Westphalian succession of the Glyn-neath area, South Wales: an internationally important geological resource. Proceedings of the Geologists' Association, 114, 291-305.

Pšenicka, J., Bek, J., Zodrow, E. L., Cleal, C. J., & Hemsley, A. 2003. A new late Westphalian fossil marattialean fern from Nova Scotia. Botanical Journal of the Linnean Society, 142, 199-212.

Cleal, C. J., Dimitrova, T. Kh. & Zodrow, E. L. 2003. Macrofloral and palynological criteria for recognising the Westphalian - Stephanian boundary. Newsletters on Stratigraphy, 39, 181-208.

Cleal, C. J. & Shute, C. H. 2003. Systematics of the Late Carboniferous medullosalean pteridosperm Laveineopteris and its associated Cyclopteris leaves. Palaeontology, 46, 353-411.

Šimunek, Z. & Cleal, C. J. 2004. Small-pinnuled odontopterid medullosaleans from the middle and upper Stephanian of Central Bohemia and Saar-Lorraine. Review of Palaeobotany and Palynology, 129, 21-38.

Cleal, C. J. & Thomas, B. A. 2004. Late Carboniferous palaeobotany of the upper Bideford Formation, north Devon: a coastal setting for a Coal Measures flora. Proceedings of the Geologists' Association, 115, 267-281.

Cleal, C. J. 2004. IGCP 469 Late Westphalian terrestrial biotas and environments of the Variscan Foreland and adjacent intramontane basins. Geologica Balcanica, 34, 3-10.

Cleal, C. J., Tenchov, Y. & Zodrow, E. L. 2004. Review of the late Westphalian early Stephanian macrofloras of the Dobrudzha Coalfield, Bulgaria. Geologica Balcanica, 34, 11-20.

Cleal, C. J. & Thomas, B. A. 2005. Palaeozoic tropical rainforests and their effect on global climates: is the past the key to the present? Geobiology, 3, 13-31.

Cleal, C. J. 2005. The Westphalian macrofloral record from the cratonic central Pennines Basin, UK. Zeitschrift der Deutschen Gesellschaft für Geowissenschaften, 156, 387-410.

Cleal, C. J., Lazarus, M. & Townsend, A. 2005. Illustrations and illustrators during the 'Golden Age' of palaeobotany: 1800-1840. In: Bowden, A. J., Burek, C. V. & Wilding, R (ed.) History of palaeobotany: selected essays. Geological Society of London, Special Publication, 241, 41-61.

Burek, C. V. & Cleal, C. J. 2005. The life and work of Emily Dix (1904-1972). In: Bowden, A. J., Burek, C. V. & Wilding, R (ed.) History of palaeobotany: selected essays. Geological Society of London, Special Publication, 241, 181-196.

Dimitrova, T. Kh., Cleal, C. J. & Thomas, B. A. 2005. Palynology of late Westphalian early Stephanian coal-bearing deposits in the eastern South Wales Coalfield. Geological Magazine, 142, 809-821.

Zodrow, E. L., Cleal, C. J. & Tenchov, Y. G. 2006. The Pennsylvanian-aged Linopteris obliqua tree from the Sydney Coalfield, Nova Scotia. Canadian Palaeobiology, 13, 5-8.

Zodrow, E. L., Šimunek, Z., Cleal, C. J., Bek, J. & Pšenicka, J. 2006. Taxonomic revision of the Palaeozoic marattialean fern Acitheca. Review of Palaeobotany and Palynology, 138, 239-280.

Cleal, C. J. & Shute, C. H. 2007. The effect of drying on epidermal cell parameters preserved on plant cuticles. Acta Palaeobotanica, 47, 315-326.
Zodrow, E. L., Tenchov, E. L. & Cleal, C. J. 2007. The arborescent Linopteris obliqua plant (Medullosales, Pennsylvanian). Bulletin of Geosciences, 82, 51-84.

Dimitrova, T. Kh. & Cleal, C. J. 2007. Palynological evidence for late Westphalian–early Stephanian vegetation change in the Dobrudzha Coalfield, NE Bulgaria. Geological Magazine, 144, 513-524.

Cleal, C. J., Tenchov, Y. G., Dimitrova, T. Kh., Thomas, B. A. & Zodrow, E. L. 2007. Late Westphalian-Early Stephanian vegetational changes across the Variscan Foreland. In Wong, Th. E (ed.) Proceedings of the XVth International Congress on Carboniferous and Permian Stratigraphy. Utrecht, the Netherlands, 10-16 August 2003. Royal Netherlands Academy of Arts and Sciences, Amsterdam, pp. 367-377.

Cleal, C. J., Zodrow, E. L. & Šimunek, Z.. 2007. Leaf cuticles from the Pennsylvanian-aged medullosalean Odontopteris cantabrica Wagner. Acta Palaeobotanica, 47, 327-337.

Cleal, C. J. 2007. The Westphalian-Stephanian macrofloral record from the South Wales Coalfield. Geological Magazine, 144, 465-486.

Opluštil, S. & Cleal, C. J. 2007. A comparative analysis of some Late Carboniferous basins of Variscan Europe. Geological Magazine, 144, 417-448.

Hilton, J. & Cleal C. J. 2007. The relationship between Euramerican and Cathaysian tropical floras in the Late Palaeozoic: palaeobiogeographical and palaeogeographical implications. Earth-Science Reviews, 85, 85-116.

Fraser, H. E. & Cleal, C. J. 2007. The contributions of British women to Carboniferous palaeobotany during the first half of the 20th century. In: Burek, C. V. & Higgs B (ed.) The role of women in the history of geology. Geological Society of London, Special Publication, 281, 51-82.

Cleal, C. J. 2008. IGCP 469 Late Variscan terrestrial biotas and palaeoenvironments. Studia Geologica Polonica, 129, 7-8.

Cleal, C. J. 2008. Macrofloral biostratigraphy of the Ottweiler Group in Saar-Lorraine and its consequences for Stephanian palynostratigraphy and geochronology. Studia Geologica lPolonica, 129, 9-23.

Cleal, C. J. 2008. Palaeofloristics of Middle Pennsylvanian lyginopteridaleans in Variscan Euramerica. Palaeogeography, Palaeoclimatology, Palaeoecology, 261, 1-14.

Cleal, C. J. 2008. Westphalian-Stephanian macrofloras of the southern Pennines Basin, UK. Studia Geologica Polonica, 129, 25-41.

Cleal, C. J. 2008. Palaeofloristics of Middle Pennsylvanian medullosaleans in Variscan Euramerica. Palaeogeography, Palaeoclimatology, Palaeoecology, 268, 164-180.

Pšenicka, J., Bek, J., Cleal, C. J., Wittry, J. & Zodrow, E. L. 2009. Description of synangia and spores of the holotype of the Carboniferous fern Lobatopteris miltoni, with taxonomic comments. Review of Palaeobotany and Palynology, 155, 133-144.

Cleal, C. J., Fraser, H. E., Lazarus, M. & Dannell, G. 2009. The forests before the flood: the palaeobotanical contributions of Edmund Tyrell Artis (1789-1847). Earth Sciences History, 28, 245-275.

Cleal, C. J., Shute, C. H., Hilton, J. & Carter, J. 2009. A revision of the Pennsylvanian-aged Eremopteris-bearing seed plant. International Journal of Plant Sciences, 170, 666-698.

Cleal, C. J. 2010. Proposal to conserve the name Eusphenopteris Gothan ex Simson-Scharold against Eusphenopteris Kidst (fossil Lyginopteridales). Taxon, 59, 653-655.

Tenchov, Y. G. & Cleal, C. J. 2010. Neuralethopteris foliage (Medullosales) in the Carboniferous of Dobrudzha Coalfield, Bulgaria. Review of Palaeobotany and Palynology, 158, 298-307.

Cleal, C. J. & Thomas, B. A. 2010. Botanical nomenclature and plant fossils. Taxon, 59, 261-268.

Cleal, C. J. & Thomas, B. A. 2010. Proposals to modify the provisions in the Code for naming plant fossils. Taxon, 59, 312-313.

Thomas, B. A., Zodrow, E. L. & Cleal, C. J. 2010. Leafy branches of Bothrodendron punctatum from the Westphalian D (Asturian) of Nova Scotia, Canada. Atlantic Geology, 46, 1-6.

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